Chapter 10 · Plant Anatomy · 3 questions in the paper · 27 min read

Plant Anatomy

What the syllabus asks for 3
  • Plant tissues: characters, classification, structure and functions
  • Types of vascular bundles
  • T.S. of monocot and dicot root, stem and leaf

Plant Anatomy carries about 3 of the 40 Botany questions, and it is the most diagram-driven chapter of the paper: the setter describes (or draws) a transverse section and asks what organ, what plant group, or what tissue you are looking at. The content falls into three blocks — the tissues themselves (meristems and the simple and complex permanent tissues), the vocabulary of vascular bundles, and the six standard sections: dicot and monocot root, stem and leaf. Master the small set of diagnostic features that separate those six slides and this chapter is free marks.

What the paper asks from this chapter
  • Meristems — characters (dense cytoplasm, thin walls, no big vacuole); apical (length), intercalary (grasses), lateral (thickness: vascular and cork cambium); primary vs secondary (cork cambium by dedifferentiation); protoderm, procambium, ground meristem.
  • Simple tissues — parenchyma (chlorenchyma, aerenchyma), collenchyma (living, pectin corners, flexible), sclerenchyma (dead, lignified; fibres and sclereids/stone cells).
  • Complex tissues — xylem (tracheids, vessels, fibres dead; parenchyma living) and phloem (enucleate sieve tubes + companion cells, bast fibres, parenchyma); proto-/metaxylem; exarch root, endarch stem.
  • Bundle words — radial (roots); conjoint collateral; bicollateral (Cucurbitaceae); amphivasal = xylem outside (Dracaena); amphicribral = phloem outside (ferns); open (cambium present) vs closed.
  • Root sections — epiblema with root hairs, endodermis with Casparian strips and passage cells, radial exarch bundles; dicot diarch–hexarch, pith absent; monocot polyarch, big pith.
  • Stem sections — dicot: ring of open bundles, collenchyma hypodermis, medullary rays; monocot: scattered closed bundles with sclerenchyma sheath, lysigenous cavity, undifferentiated ground tissue.
  • Leaf sections — dorsiventral dicot (palisade above, spongy below, hypostomatic) vs isobilateral monocot (uniform mesophyll, bulliform/motor cells, dumb-bell guard cells).

10.1 Plant tissues: characters, classification, structure and functions

A tissue is a group of cells of common origin working together. Plant tissues divide first by one question — can the cells still divide? If yes, they are meristematic; if they have matured and stopped, they are permanent.

Meristematic tissues

Meristem cells are small and isodiametric, with thin cellulose walls, dense cytoplasm, a large central nucleus and no large vacuole, and they are packed together without intercellular spaces. They divide repeatedly by mitosis. Three classifications are asked:

By position. - Apical meristem — at root and shoot tips; causes primary growth in length. - Intercalary meristem — pieces of apical meristem left at the bases of internodes and leaves, famous in grasses (it is why a lawn regrows after mowing and grazing). - Lateral meristem — cylinders along the axis, the vascular cambium and cork cambium (phellogen); causes growth in thickness (secondary growth) in dicots and gymnosperms. Monocots generally lack lateral meristems.

By origin. The promeristem is the few initials at the very tip; primary meristems descend from them directly (apical, intercalary and the intrafascicular cambium); secondary meristems arise later when permanent cells regain the power to divide — dedifferentiation — as the cork cambium and interfascicular cambium do.

By function (Haberlandt). At an apex, protoderm makes the epidermis, procambium makes primary xylem, phloem and vascular cambium, and ground meristem makes cortex, endodermis, pericycle, pith and medullary rays.

Simple permanent tissues

Permanent cells arise from meristems by differentiation. Simple tissues are built of one cell type.

photograph
1Sclerenchyma — thick, lignified walls, narrow lumen, no living contents2Parenchyma — thin-walled, large, living cells with intercellular spaces
Figure 1. Parenchyma and sclerenchyma side by side in a transverse section of a *Cucurbita* stem (×100). The difference is in the wall: thin cellulose and large living cells in parenchyma; thick, uniformly lignified walls and a tiny lumen in sclerenchyma, whose cells are dead at maturity. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

Tissue Cells Walls Function and notes
Parenchyma living, isodiametric thin, cellulose the packing/storage tissue; with chloroplasts = chlorenchyma (photosynthesis); with big air spaces = aerenchyma (hydrophytes, buoyancy and aeration)
Collenchyma living, elongated unevenly thickened with pectin (angular, lamellar, lacunar types) flexible mechanical support in still-growing organs — petioles, young stems, leaf midribs
Sclerenchyma dead at maturity thick, uniformly lignified, simple pits, narrow lumen rigid mechanical support; long fibres (bundles) and short sclereids (stone cells) — grit in pear, hard seed coats and nutshells

The collenchyma–sclerenchyma contrast is a perennial MCQ: collenchyma is living and plastic (grows with the organ), sclerenchyma is dead and rigid (only in organs that have finished elongating).

Complex permanent tissues

Xylem carries water and minerals upwards from root to leaf.

Element Living? Structure and function
Tracheids dead elongated and tapering, lignified, water passes through bordered pits; the only conducting element of pteridophytes and gymnosperms; conduction + support
Vessels dead rows of cells with perforated end walls forming continuous wide pipes; the efficient conductors characteristic of angiosperms
Xylem fibres dead sclerenchymatous; support
Xylem parenchyma living storage and sideways conduction — the only living xylem element

Phloem translocates prepared food (mainly sucrose) from source to sink, in either direction as needed.

photograph
1Vessel with pitted, lignified wall — the lumen is empty (dead cell)2Vessel with ladder-like (scalariform) thickening of the wall
Figure 2. Xylem vessels in longitudinal section (*Cucurbita* stem, ×400). The walls are lignified and carry pits and bars of thickening; the cross-walls between vessel elements have broken down, so the vessel is an open pipe made of dead cells. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

photograph
1Sclereid — thick layered wall with simple pits, almost no lumen
Figure 3. Sclereids (stone cells) in the flesh of a pear (×400). A cluster of isodiametric cells with very thick, layered, lignified walls and a narrow lumen gives pear its gritty texture. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

Element Living? Structure and function
Sieve tube elements living but enucleate end-to-end tubes joined by perforated sieve plates; the conducting channel
Companion cells living narrow, dense cytoplasm, large nucleus; keep the sieve tube alive and load its sugar; confined to angiosperms
Phloem fibres (bast fibres) dead sclerenchymatous; support — jute, flax and hemp of commerce
Phloem parenchyma living storage; absent in most monocots — a standard named exception

Xylem formed first (protoxylem, narrow annular/spiral elements that can stretch) differs from later metaxylem (wide, pitted). Their arrangement names the organ: exarch — protoxylem towards the periphery — in roots; endarch — protoxylem towards the centre — in stems. ("Roots are EXtroverts, stems are ENDoverts.")

Special secretory tissues — laticifers carrying latex (Euphorbia, Ficus, rubber), resin ducts (pines), oil glands and nectaries — complete the list.

photograph
1Sieve tube element — thin-walled, living, enucleate
Figure 4. Phloem sieve tubes in longitudinal section (*Cucurbita* stem, ×400). Sieve tubes are living cells joined end to end; their thin cross-walls are perforated as sieve plates and they have lost their nucleus, being kept alive by the companion cell beside each one. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

figure
Figure 5. Water's inward road through a root in four doors: the epiblema and its hairs gather it, the cortex stores and passes it, the endodermis with its Casparian strips forces it through living membranes, and behind the pericycle — birthplace of lateral roots — the xylem vessels lift it up the plant.

Reading four unknown slides

Problem. Four unlabeled transverse sections show: (i) bundles in a ring, each with cambium, xylem endarch, a starch sheath outside the bundles; (ii) bundles scattered in ground tissue, each closed with a sclerenchyma sheath; (iii) radial bundles with four xylem arms, a small pith, pericycle present; (iv) polyarch radial bundles around a large pith. Identify each. Solution. (i) Dicot stem — ring, open bundles, endarch xylem and the starch sheath are its signature. (ii) Monocot stem — scattered, closed, sheathed bundles in undifferentiated ground tissue. (iii) Dicot root — radial bundles, tetrarch (four arms), small pith; the cambium for its secondary growth arrives later. (iv) Monocot root — polyarch xylem and a large pith, no cambium ever. Four slides, four answers, each from two or three diagnostic features — exactly how the paper frames it.

10.2 Types of vascular bundles

A vascular bundle is a strand of xylem and phloem. The paper asks you to name bundles, so learn the vocabulary exactly:

  • Radial — xylem and phloem in separate groups on different radii, alternating. The root arrangement.
  • Conjoint — xylem and phloem together in one bundle (stems and leaves):
  • Collateral — phloem on the outer side only, xylem inside (the usual stem bundle).
  • Bicollateral — phloem on both sides of the xylem (Cucurbitaceae — Cucurbita, Luffa; also Solanum).
  • Concentric — one tissue completely surrounds the other:
  • Amphivasal (leptocentric) — xylem surrounds phloem (Dracaena, Yucca).
  • Amphicribral (hadrocentric) — phloem surrounds xylem (ferns).
  • Open — cambium present between xylem and phloem: secondary growth possible (dicot stems).
  • Closed — no cambium: no secondary growth (monocot stems, all leaves).
The one-syllable trap

Amphi-VAS-al: vascular tissue — wood, xylem — outside. Amphi-CRIB-ral: cribrum = a sieve — sieve tubes, phloem — outside.

figure
Figure 6. The named bundle types. Blue is xylem, green is phloem, and the thin gold strip in the collateral bundle is the cambium that makes it 'open'.

10.3 T.S. of monocot and dicot root, stem and leaf

Roots — the shared ground plan

Work from outside in. Epiblema (rhizodermis): one layer of thin-walled cells, many extended into root hairs; no cuticle, no stomata — it absorbs. Cortex: broad parenchyma with air spaces. Endodermis: one ring of barrel-shaped cells whose radial and transverse walls carry a waterproof band of suberin and lignin, the Casparian strip, forcing water and minerals to cross a living membrane (the cell checks what enters the stele). A few endodermal cells opposite the protoxylem stay thin-walled — passage cells, letting water through quickly. Pericycle: one layer inside the endodermis; in roots it is the layer that gives rise to lateral roots (endogenous origin). Stele: xylem and phloem in separate alternating radial groups, xylem exarch, with conjunctive parenchyma between them.

photograph
1Epiblema (piliferous layer) — one cell thick, no cuticle, bears root hairs2Cortex — a wide zone of thin-walled parenchyma packed with starch grains3Endodermis — innermost cortical layer, its walls carrying Casparian strips4Xylem of the stele — exarch, here tetrarch; the box outlines the whole stele
Figure 7. Transverse section of a young dicot root (*Ranunculus*), ×100. The boxed stele is tiny compared with the cortex — that proportion is itself diagnostic of a dicot root. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

Feature Dicot root (e.g. Ranunculus, gram) Monocot root (e.g. maize)
Xylem groups 2–6 — diarch, triarch, tetrarch … hexarch many, more than 6 — polyarch
Pith absent (xylem meets at the centre) large, well-developed parenchyma
Secondary growth occurs (cambium forms) does not occur
Shared signs both: root hairs, Casparian strips and passage cells, radial and exarch bundles

Stems

Dicot stem (Helianthus). Epidermis with cuticle, stomata and multicellular hairs; hypodermis of collenchyma; general cortex; endodermis full of starch grains (the starch sheath); pericycle with semilunar sclerenchyma caps over the bundles; then a single ring of vascular bundles (a eustele), uniform in size, each conjoint, collateral, open and endarch, separated by medullary rays; large central pith. The open bundles and ring layout are exactly what secondary growth needs: the cambium completes itself into a ring.

photograph
1Epidermis, drawn out into ridges and carrying a cuticle2Hypodermis of collenchyma, thickest under each ridge, then a narrow cortex3Sclerenchymatous cap (pericyclic fibres) standing over each bundle4Vascular bundle — conjoint, collateral and open; the wide cells are metaxylem5Pith of parenchyma; medullary rays run out between the bundles
Figure 8. Transverse section of a young dicot stem (*Helianthus*), ×40. The vascular bundles lie in a single ring around a wide pith. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

Monocot stem (Zea mays). Epidermis with thick cuticle; hypodermis of sclerenchyma; inside it, no cortex/endodermis/pith at all — one continuous parenchymatous ground tissue; vascular bundles scattered through it (an atactostele), numerous, smaller and denser towards the periphery, each conjoint, collateral and closed, wrapped in a sclerenchymatous bundle sheath, and carrying a water-filled lysigenous cavity (protoxylem lacuna) where the earliest protoxylem vessel broke down. Scattered closed bundles are why a maize stem can never thicken into a trunk.

photograph
1Epidermis, with the remains of root hairs2Cortex — wide and parenchymatous, as in a dicot root3Ring of many alternating xylem and phloem groups (polyarch)4Large central pith — the feature a dicot root does not have
Figure 9. Transverse section of a monocot root (*Zea mays*), ×40. The vascular bundles form a ring enclosing a large central pith. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

Leaves

A horizontal dicot leaf is two-faced — dorsiventral. The upper epidermis has a thick cuticle and few or no stomata; the lower has many stomata (hypostomatic). The mesophyll is differentiated: palisade parenchyma above — elongated cells crammed with chloroplasts in the best light, the main photosynthetic layer — and spongy parenchyma below, loose cells and air spaces leading to the stomata. Vein bundles are conjoint, collateral and closed, with xylem towards the upper side and phloem towards the lower.

photograph
1Upper epidermis with a thick cuticle and few or no stomata2Palisade parenchyma — elongated cells on end, holding most of the chloroplasts3Spongy mesophyll — loose cells with large air spaces for gas exchange4Lower epidermis — thinner cuticle, most of the stomata5Midrib vascular bundle — xylem above, phloem below
Figure 10. Transverse section of a dorsiventral dicot leaf (*Ligustrum*, privet), ×100. Palisade cells stand on end directly under the upper epidermis; the spongy mesophyll below is full of air spaces. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

photograph
1Epidermis with a thick cuticle, underlain by a sclerenchymatous hypodermis2Small, crowded peripheral bundles3Large, widely spaced central bundle — conjoint, collateral and closed4Continuous parenchymatous ground tissue (no cortex, endodermis or pith)
Figure 11. Transverse section of a monocot stem (*Zea mays*), ×40. The bundles are scattered through a continuous ground tissue — small and crowded near the surface, large and widely spaced towards the centre. Source: Berkshire Community College Bioscience Image Library / Wikimedia Commons (CC0)

A vertical monocot (grass/maize) leaf is alike on both sides — isobilateral: stomata on both epidermes with dumb-bell-shaped guard cells (dicot guard cells are kidney-shaped); bulliform (motor) cells — big, thin-walled, watery cells in the upper epidermis that lose turgor in dry weather and roll the leaf inwards to cut water loss; mesophyll undifferentiated (no palisade/spongy split); bundles in a parallel row between the veins.

figure
Figure 12. Ground plans of the two stem sections. Left, dicot: one ring of similar open bundles around a pith. Right, monocot: closed bundles scattered in ground tissue, smaller and denser towards the outside, with no distinct pith.

Worked example — naming an unknown slide

Problem. A stained transverse section shows: a cuticularised epidermis with stomata and hairs; a collenchymatous hypodermis; conjoint, collateral, endarch vascular bundles of nearly equal size arranged in one ring, each with cambium; medullary rays between the bundles; and a large central pith. Identify the section.

Solution. Eliminate in order. Root hairs and radial, exarch bundles are absent, and a cuticle with stomata is present — so it is not a root. The bundles are in a ring, not scattered — so it is not a monocot stem. Open bundles (cambium present), collenchyma hypodermis, medullary rays and a large pith complete the picture: a young dicot stem. Each decision used one feature the alternatives cannot have — that is how these marks are earned quickly.

Reading an unknown slide in ten seconds

Root or stem? Radial bundles + exarch xylem + root hairs = root; conjoint bundles = stem. Dicot or monocot root? Count the xylem groups and look for a pith. Dicot or monocot stem? Ring = dicot, scattered = monocot — then confirm with open vs closed. Leaf? Palisade present = dicot; bulliform cells and dumb-bell guard cells = grass.

figure
Figure 13. The tissue teams in four drawers: the three simple permanent tissues by wall and fate, the water-conducting xylem, the food-conducting phloem with its companion cells, and the bundle geometries that identify every section on the slide.

Common mistake

Four pairs the paper swaps: collenchyma is living with pectin corners, sclerenchyma dead with lignin — support that flexes vs support that stiffens; xylem is exarch in the root, endarch in the stem; the endodermis of a root carries Casparian strips, the endodermis of a dicot stem is a starch sheath without them; and companion cells belong to phloem only — xylem's dead pipes need no attendants.

Distinguishing root vs stem and monocot vs dicot in T.S.

Problem. A transverse section shows radial vascular bundles with exarch protoxylem and more than six xylem patches (polyarch) surrounding a large parenchymatous pith. Identify the organ and group.

Solution. Radial bundles + exarch protoxylem = root; polyarch condition (>6 xylem arches) + large persistent pith = monocot root (e.g., maize).

Formula and fact sheet

  • Meristem characters: thin walls, dense cytoplasm, big nucleus, tiny vacuoles, no spaces. Apical = length; intercalary = grasses; lateral = thickness (vascular + cork cambium).
  • Origins: protoderm → epidermis; procambium → vascular tissues; ground meristem → cortex, pith etc. Cork cambium = secondary meristem by dedifferentiation.
  • Parenchyma: living, thin; chlorenchyma photosynthesises; aerenchyma aerates hydrophytes.
  • Collenchyma: living, pectin-thickened corners, flexible. Sclerenchyma: dead, lignified — fibres + sclereids (pear grit, seed coats).
  • Xylem: tracheids, vessels, fibres dead; parenchyma the only living element. Vessels = angiosperms; gymnosperms conduct by tracheids alone.
  • Phloem: sieve tubes enucleate + companion cells; bast fibres = jute/flax; phloem parenchyma absent in most monocots.
  • Exarch = root, endarch = stem; protoxylem narrow and early, metaxylem wide and later.
  • Bundles: radial (root); collateral; bicollateral (Cucurbitaceae); amphivasal — xylem outside (Dracaena); amphicribral — phloem outside (ferns); open = cambium, closed = none.
  • Root: epiblema + root hairs; Casparian strip in endodermis; passage cells opposite protoxylem; lateral roots from pericycle.
  • Dicot root: 2–6 xylem groups, pith absent. Monocot root: polyarch, big pith.
  • Dicot stem: collenchyma hypodermis, starch-sheath endodermis, ring of open bundles, medullary rays, pith. Monocot stem: sclerenchyma hypodermis, ground tissue, scattered closed bundles + sheath + lysigenous cavity.
  • Leaves: dorsiventral — palisade up, spongy down, hypostomatic, kidney guard cells; isobilateral — uniform mesophyll, bulliform cells, dumb-bell guard cells, stomata both sides.

Rapid-revision tables and high-yield facts

Feature Diagnostic
Meristems apical (length), intercalary (grass regrowth), lateral (girth)
Collenchyma / sclerenchyma pectin corners, living / lignin, dead
Xylem conductors tracheids (pits), vessels (perforated tubes)
Phloem sieve tubes + companion cells; bidirectional
Root bundle radial, exarch; dicot 2–6 arms, monocot polyarch
Stem bundle conjoint collateral; dicot open in ring, monocot closed scattered
Leaf dicot dorsiventral, monocot isobilateral + bulliform cells
C₄ leaf Kranz bundle sheath with chloroplasts
Casparian strip endodermis of root (suberin)
Lateral roots arise from pericycle

Chapter summary

  • Tissues are meristematic (still dividing: apical, intercalary and lateral; primary and, by dedifferentiation, secondary) or permanent (simple and complex).
  • Parenchyma packs and stores (aerenchyma in hydrophytes), living pectin-rich collenchyma supports growing organs flexibly, and dead lignified sclerenchyma (fibres and stone cells) supports finished ones rigidly.
  • Xylem moves water up through dead tracheids and vessels, its parenchyma alone alive; phloem moves food through enucleate sieve tubes serviced by companion cells, with bast fibres for strength.
  • Bundle names encode geometry: radial in roots; collateral and bicollateral conjoint bundles in stems; concentric amphivasal (xylem out) and amphicribral (phloem out); open bundles carry cambium, closed do not.
  • Roots show epiblema with hairs, a Casparian-striped endodermis with passage cells, pericycle that births lateral roots, and exarch radial groups — few and pith-less in dicots, polyarch with a big pith in monocots.
  • Dicot stems ring their open bundles round a pith (ready for secondary growth); monocot stems scatter closed, sheathed bundles with a lysigenous cavity through undifferentiated ground tissue.
  • Dorsiventral dicot leaves split mesophyll into palisade and spongy layers and hide stomata below; isobilateral grass leaves keep mesophyll uniform, bear stomata and dumb-bell guard cells on both faces, and roll up by their bulliform cells.

Practice questions

Set A — Recall (25 questions)

  1. Cells of a meristem are characterised byathin walls, dense cytoplasm and a large nucleusbthick lignified wallsca huge central vacuoledabsence of a nucleus
  2. Primary growth in the length of root and shoot is caused by theavascular cambiumbcork cambiumcintercalary meristem onlydapical meristem
  3. Intercalary meristems, found at the bases of internodes, are characteristic ofagrassesbmango treescfernsdmushrooms
  4. Which of the following is a secondary meristem?acork cambium (phellogen)broot apical meristemcshoot apical meristemdprotoderm
  5. The tissue that forms the basic packing and storage tissue of plants isasclerenchymabcollenchymacparenchymadxylem
  6. Aerenchyma, parenchyma with large air spaces, is typically found inaxerophytesbmesophytescparasitesdhydrophytes
  7. Collenchyma differs from sclerenchyma in beingadead and lignifiedbwithout wallsca conducting tissuedliving with pectin-thickened walls
  8. Sclerenchyma cells arealiving and thin-walledbdead with thick lignified wallscenucleate and livingdfull of chloroplasts
  9. Stone cells responsible for the gritty texture of pear fruit areafibresbtracheidsccompanion cellsdsclereids
  10. Wide vessels with perforated end walls are characteristic of the xylem ofapteridophytesbgymnospermscangiospermsdbryophytes
  11. The only living element of xylem is theavesselbxylem parenchymactracheiddxylem fibre
  12. A sieve tube element at maturity lacks aanucleusbcell wallccytoplasmdsieve plate
  13. The jute and flax fibres of commerce areaxylem fibresbbast (phloem) fibrescsurface fibresdsclereids
  14. Phloem parenchyma is absent in mostamonocotsbdicotscgymnospermsdferns
  15. Exarch xylem (protoxylem towards the periphery) is found in theastembleafcrootdflower
  16. Bicollateral vascular bundles are characteristic of the familyaPoaceaebCucurbitaceaecLiliaceaedBrassicaceae
  17. In an amphivasal (leptocentric) bundle, as in Dracaena,aphloem surrounds the xylembxylem and phloem alternateccambium surrounds bothdxylem surrounds the phloem
  18. Radial vascular bundles, with xylem and phloem on different radii, occur inarootsbdicot stemscmonocot stemsdleaves
  19. A vascular bundle is called open when it possessesacambium between xylem and phloemba bundle sheathca lysigenous cavitydphloem on both sides
  20. Casparian strips are found in the cells of theaepiblemabpericyclecendodermisdhypodermis
  21. A well-developed pith is absent, the xylem meeting at the centre, in theamonocot rootbdicot rootcdicot stemdmonocot stem
  22. In a monocot stem the vascular bundles areain a single ring and openbscattered and closedcradial and exarchdbicollateral
  23. A lysigenous (water-containing) cavity is a feature of the vascular bundles ofasunflower stembgram rootcmaize stemdmango leaf
  24. Bulliform (motor) cells that roll the leaf in dry weather occur in the leaves ofamangobpeacgrassesdCucurbita
  25. In a dorsiventral (dicot) leaf, the mesophyll is differentiated intoacortex and stelebxylem and phloemcpalisade and bulliform tissuedpalisade and spongy parenchyma
Answers and explanations — Set A

1. (a) — Meristem cells are small, thin-walled and crowded with cytoplasm and nucleus; big vacuoles appear only as cells mature.

2. (d) — Apical meristems at the tips lengthen the axis; lateral meristems thicken it.

3. (a) — Grass nodes keep intercalary meristems, so lawns regrow after cutting.

4. (a) — The phellogen arises from permanent cortical cells that dedifferentiate — the definition of a secondary meristem.

5. (c) — Parenchyma fills, stores and (as chlorenchyma) photosynthesises.

6. (d) — Air cavities give hydrophytes buoyancy and a path for oxygen.

7. (d) — Collenchyma stays alive with plastic pectin thickenings and can stretch; sclerenchyma is dead and lignified.

8. (b) — Maturity brings a thick lignified wall, simple pits and loss of the protoplast.

9. (d) — Short, heavily thickened sclereids make pear gritty and seed coats hard.

10. (c) — True vessels are the angiosperm speciality; gymnosperms and ferns manage with tracheids.

11. (b) — Vessels, tracheids and fibres are all dead pipes and props; only xylem parenchyma lives.

12. (a) — The nucleus disintegrates; the companion cell's nucleus runs the sieve tube instead.

13. (b) — Bast fibres are the sclerenchyma of the phloem — jute, flax, hemp.

14. (a) — A standard named absence: monocot phloem usually lacks parenchyma.

15. (c) — Roots are exarch; stems are endarch.

16. (b) — Cucurbita and its relatives carry phloem on both faces of the xylem.

17. (d) — Amphivasal: vascular (xylem) tissue outside, phloem in the middle.

18. (a) — Alternate xylem and phloem poles on different radii diagnose a root.

19. (a) — "Open for (secondary) growth": the fascicular cambium is present.

20. (c) — Suberin–lignin bands in endodermal walls seal the stele against uncontrolled entry.

21. (b) — In dicot roots the few xylem arms meet in the middle; monocot roots keep a large pith.

22. (b) — Atactostele: bundles scattered in ground tissue, all closed.

23. (c) — The earliest protoxylem of the maize bundle breaks down into a water-filled lacuna.

24. (c) — Bulliform cells in the upper epidermis of grass leaves roll them against drought.

25. (d) — Palisade above for light capture, spongy below for gas exchange.

Set B — Understanding (15 questions)

  1. Meristematic cells lack a large central vacuole mainly because they areadead cellsbstoring foodcdividing repeatedly and packed with cytoplasmdlignified
  2. Collenchyma, rather than sclerenchyma, supports growing petioles because it isafull of air spacesbdead and rigidcliving and able to stretch as the organ growsdmade only of pectin-free walls
  3. Vessels conduct water more efficiently than tracheids because theiraend walls are perforated, forming continuous wide pipesbwalls are unlignifiedccells retain nucleidpits are bordered
  4. A sieve tube depends on its companion cell because the sieve tubeahas no wallbis deadchas lost its nucleus and most organellesdcannot make sugar flow downwards
  5. A closed vascular bundle cannot produce secondary tissues because it lacksaxylem vesselsbcambiumcphloem fibresda bundle sheath
  6. A section with radial bundles and exarch xylem must be a root, because in stems the bundles areaconjoint and the xylem endarchbabsentcalways bicollateraldwithout phloem
  7. Passage cells in the endodermis are important because they allowaeasy entry of water into the xylem opposite the protoxylembphotosynthesis in the rootcformation of root hairsdstorage of starch
  8. The ring arrangement of bundles in a dicot stem makes secondary growth possible because the bundle cambiaaare deadblie scatteredccan be joined by interfascicular cambium into a complete ringdproduce only phloem
  9. In a maize stem the vascular bundles are smaller and more crowded near the periphery, which is whereamechanical stress in bending is greatestbphotosynthesis is fastestcthe pith is largestdroot hairs form
  10. When bulliform cells lose water and become flaccid, the grass leafaopens flatbrolls inwards, reducing the exposed surface and transpirationcsheds its mesophylldcloses its stomata directly
  11. A hypostomatic leaf (stomata mostly on the lower surface) is an adaptation toaincrease photosynthesisbabsorb more CO₂cattract pollinatorsdreduce water loss
  12. Sclerenchyma is absent from the growing tip region because its dead, lignified cellsaconduct waterbdivide too fastcphotosynthesisedcannot stretch as the organ elongates
  13. Gymnosperms conduct water less efficiently than angiosperms because their xylem containsaonly vesselsbonly tracheids, water crossing end walls through pitsccompanion cellsdno lignin
  14. Palisade tissue lies just below the upper epidermis of a dorsiventral leaf becauseathe lower side has more lightbstomata are absent abovecit stores waterdthe upper side receives most light for photosynthesis
  15. The innermost cortical layer of a dicot stem is called the starch sheath because it is homologous to the endodermis and its cells are loaded withaoil dropsblatexccrystalsdstarch grains
Answers and explanations — Set B

1. (c) — Continual division keeps meristem cells small, dense and vacuole-poor; vacuolation comes with maturation.

2. (c) — Living pectin walls are plastic: support without imprisoning growth.

3. (a) — Perforations remove end-wall resistance; tracheid water must seep pit to pit.

4. (c) — Enucleate sieve elements are kept alive, and loaded, by their nucleate companions.

5. (b) — No fascicular cambium, no secondary xylem or phloem.

6. (a) — Stem bundles are conjoint with protoxylem towards the centre (endarch); the root pattern is the reverse.

7. (a) — Thin-walled passage cells short-circuit the Casparian seal right where the xylem poles lie.

8. (c) — Medullary-ray cells between bundles dedifferentiate into interfascicular cambium, closing the ring.

9. (a) — Sheathed peripheral bundles act like the steel near the surface of a beam.

10. (b) — The motor cells hinge the blade shut around its stomata in drought.

11. (d) — Shaded lower stomata lose less water than sun-struck upper ones would.

12. (d) — Rigid dead tissue in an elongating zone would tear or block extension.

13. (b) — Without vessels every water molecule must cross pit membranes between tracheids.

14. (d) — Chloroplast-packed columns are placed where light actually arrives.

15. (d) — The stem "endodermis" is recognised by its abundant starch rather than Casparian strips.

Set C — Application (10 questions)

  1. A stem section shows numerous closed bundles, each with a sclerenchymatous sheath and a water cavity, scattered in ground tissue. It is aadicot stembdicot rootcmonocot rootdmonocot stem
  2. A root section shows radial bundles with four xylem groups meeting at the centre and no pith. It is aamonocot rootbdicot rootcmonocot stemddicot stem
  3. A root section shows many (more than six) alternating xylem and phloem groups around a large central pith. It is aadicot rootbdicot stemcleafdmonocot root
  4. A stem section shows bicollateral bundles in two rings. The plant most likely belongs toaPoaceaebCucurbitaceaecLiliaceaedPinaceae
  5. A leaf section shows undifferentiated mesophyll, stomata on both surfaces and dumb-bell-shaped guard cells. It is aadicot leafbpine needlecfern fronddmonocot (grass) leaf
  6. The gritty particles felt while eating a pear are clusters ofaparenchymabcollenchymacsclereidsdvessels
  7. The commercial jute fibre is extracted from which tissue of the stem?abast (secondary phloem) fibresbxylem fibrescepidermal hairsdcortical parenchyma
  8. On a hot dry afternoon the leaves of grass roll into tubes. The cells mainly responsible are theapalisade cellsbguard cellscbulliform cellsdcompanion cells
  9. Root hairs, which absorb water and minerals, develop as outgrowths of theacortexbepiblemacendodermisdpericycle
  10. A bundle in which phloem occurs on both the outer and the inner side of the xylem is namedabicollateralbamphivasalcradialdamphicribral
Answers and explanations — Set C

1. (d) — Scattered closed bundles with sheath and lysigenous cavity: the monocot (maize) stem signature.

2. (b) — Tetrarch, pith-less: a dicot root.

3. (d) — Polyarch arrangement around a big pith: a monocot root.

4. (b) — Bicollateral bundles are the hallmark of Cucurbitaceae (Cucurbita, Luffa).

5. (d) — Isobilateral anatomy with dumb-bell guard cells belongs to grass-type monocot leaves.

6. (c) — Stone cells (sclereids) scattered in the soft parenchyma give the grit.

7. (a) — Jute is retted from the bast — the phloem fibres of Corchorus stems.

8. (c) — Flaccid bulliform (motor) cells fold the blade inwards against water loss.

9. (b) — Each root hair is a tubular extension of one epiblema (rhizodermis) cell.

10. (a) — Phloem flanking xylem on both sides defines the bicollateral conjoint bundle.